:root{--paper:#f5f1e8;--surface:#fffdf8;--ink:#182126;--muted-ink:#515b5c;--deep-plum:#28182b;--neural-dark:#120d18;--tissue-dark:#1b1320;--tissue-coral:#c86c58;--cue-amber:#e5ad54;--signal-cyan:#71bdc8;--coral-text:#9b493b;--amber-text:#765313;--cyan-text:#236b75;--interactive-link:#236b75;--soft-sage:#aab7a5;--border:#d8d1c5;--optogenetic-violet:#7d6ed1;--background:var(--neural-dark);--text-primary:#f2f0ec;--text-muted:#aaa6ad;--surface-raised:#17141d;--surface-warm:#1b1118;--line-subtle:#f2f0ec24;--line-strong:#f2f0ec4d;--page-gutter:clamp(1.25rem, 4vw, 4.5rem);--page-max:1380px;--header-height:5rem;--font-display:Arial, Helvetica, sans-serif;--font-body:Arial, Helvetica, sans-serif;--font-serif:Arial, Helvetica, sans-serif;--font-sans:Arial, Helvetica, sans-serif;--font-mono:Arial, Helvetica, sans-serif;--text-display-xl:clamp(4rem, 5.35vw, 5.9rem);--text-display-lg:clamp(2.75rem, 4vw, 4.25rem);--text-body-lg:clamp(1.1rem, 1.35vw, 1.2rem);--text-body:1.0625rem;--text-meta:.75rem}*{box-sizing:border-box;margin:0;padding:0}html{scroll-behavior:smooth;max-width:100%;min-height:100%;scroll-padding-top:calc(var(--header-height) + 1rem);background:var(--paper);overflow-x:clip}body{max-width:100%;min-height:100%;color:var(--ink);background:var(--paper);font-family:var(--font-sans);font-size:var(--text-body);-webkit-font-smoothing:antialiased;-moz-osx-font-smoothing:grayscale;line-height:1.68;overflow-x:clip}a{color:inherit;text-decoration:none}button,input,select,textarea{color:inherit;font:inherit}button{border:0}img,svg,video,canvas{max-width:100%;display:block}main,section,article{min-width:0}ul,ol{list-style:none}::selection{color:var(--ink);background:var(--cue-amber)}:focus-visible{outline:2px solid var(--signal-cyan);outline-offset:4px}.skip-link{z-index:1000;color:var(--neural-dark);background:var(--signal-cyan);padding:.65rem 1rem;font-weight:700;position:fixed;top:.75rem;left:.75rem;transform:translateY(-180%)}.skip-link:focus{transform:translateY(0)}@media (max-width:700px){:root{--header-height:4.5rem}}@media (prefers-reduced-motion:reduce){html{scroll-behavior:auto}*,:before,:after{scroll-behavior:auto!important;transition-duration:.01ms!important;animation-duration:.01ms!important;animation-iteration-count:1!important}}